Loffler, Friedrich August Johannes (1852-1915), a German bacteriologist. In 1882, Loffler discovered the bacilli causing glanders and swine erysipelas. In 1884, he isolated the Klebs-Loffler bacIllus, which is responsible for diptheria. He traced the cause of foot-and-mouth disease to a virus in 1893, and later developed a protective serum. Loffler attended the universities of Worzburg and Berlin. After serving as an army surgeon, he became a professor at the University of Greifswald in 1888.